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Flora of China 4: 97–101. 1999. 1
EPHEDRACEAE
麻黄科 ma huang ke
Fu Liguo (傅立国 Fu Li-kuo)1, Yu Yongfu (于永福)2; Harald Riedl3
Shrubs, subshrubs, or herbs, dioecious, rarely monoecious; stems erect, procumbent, or occasionally climbing, jointed, much branched, photosynthetic; branchlets opposite or whorled, green, terete, longitudinally grooved. Leaves opposite or in whorls of 3, scalelike, basally ± connate and reduced to membranous sheaths, generally ephemeral, mostly not photosynthetic; resin canals absent. Cones terminal or axillary, ovoid or ellipsoid. Pollen cones solitary or clustered at nodes, each cone composed of membranous bracts arranged in 2–8 decussate pairs or whorls of 3; proximal bracts empty; each distal bract subtending a male flower composed of 2 basally fused, orbicular or obovate scales (false perianth); anthers sessile or stipitate on staminal column. Seed cones opposite or in whorls of 3 or 4 at nodes, each cone composed of overlapping bracts; bracts arranged in 2–10 decussate pairs or whorls of 3, red and fleshy at maturity (rarely brown and membranous), proximal bracts empty, most distal bracts subtending an axillary female flower composed of a pair of fused, leathery scales (false perianth) enclosing ovule with a single membranous integument prolonged into a slender, tubular micropyle. Seeds 1 or 2(or 3) per cone. Cotyledons 2. Germination epigeal. 2n = 14*, ?24*, 28*, ?36*, 56*.
One genus and ca. 40 species: arid regions of E and N Africa, Asia, Europe, North America, and South America; 14 species (two endemic) in China.
The strictly flowering plant terms flower, staminal column, and anther are used here to avoid unwieldy descriptions.
Cheng Ching-yung. 1978. Ephedraceae. In: Cheng Wan-chün & Fu Li-kuo, eds., Fl. Reipubl. Popularis Sin. 7: 468–489.
1. EPHEDRA Linnaeus, Sp. Pl. 2: 1040. 1753.
麻黄属 ma huang shu
Morphological characters and geographical distribution are the same as those of the family.
1a. Bracts of seed cones almost completely free, connate only at base, light brown and membranous at maturity ........................................................................................................................................ 1. E. przewalskii
1b. Bracts of seed cones usually connate for 1/3–5/6 their length, red and fleshy at maturity.
2a. Seeds prominently longitudinally ridged, with dense, tiny projections .............................. 3. E. rhytidosperma
2b. Seeds smooth, rarely finely longitudinally striate.
3a. Integument tube 3–5 mm, usually spirally twisted ............................................................. 2. E. intermedia
3b. Integument tube 1–2(–2.5) mm, straight, curved, or slightly twisted.
4a. Shrubs or subshrubs, usually 50–150 cm.
5a. Bracts of seed cones with margin broad, membranous, often erose; integument tube ca. 1.5 mm, slightly spirally twisted; seeds 2 or 3; subshrubs usually to 50 cm .......... 4. E. lomatolepis
5b. Bracts of seed cones with margin narrower, entire or almost so; integument tube 1–2 mm, straight or slightly curved; seeds 1 or 2; shrubs or subshrubs often more than 50 cm.
6a. Apical pair of bracts of seed cones connate for 3/4–8/9 their length; seeds finely striate dorsally ................................................................................................. 9. E. likiangensis
6b. Apical pair of bracts of seed cones connate for 1/2–2/3 their length; seeds completely smooth.
7a. Herbaceous branches virgate, often pruinose, 1–1.5 mm in diam., rigid; integument tube to 2 mm, straight or slightly curved; plants to 100 cm or more .................. 10. E. equisetina
7b. Herbaceous branches not virgate, never pruinose, 1.5–2.5(–3) mm in diam., stout; integument tube ca. 0.5 mm, straight; plants to 60 cm .................................... 5. E. saxatilis
4b. Shrubs or subshrubs not more than 40 cm.
8a. Plants (15–)20–60 cm.
9a. Integument tube ca. 0.5 mm, straight; bracts of seed cones in 2 or 3 pairs .....
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